And it is green. Its main functions are photosynthesis



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Mesophyll[change | change source]
Most of the interior of the leaf between the upper and lower layers of epidermis is a tissue called the mesophyll (Greek for "middle leaf"). This assimilation tissue is the main place photosynthesis takes place in the plant. The products of photosynthesis are sugars, which can be turned into other products in plant cells.
In ferns and most flowering plants, the mesophyll is divided into two layers:

  • An upper palisade layer of tightly packed, vertical cells, one to two cells thick. Its cells contain many chloroplasts. The chloroplasts move by a process called "cyclosis". The slight separation of the cells provides maximum absorption of carbon dioxide. Sun leaves have a multi-layered palisade layer, while shade leaves closer to the soil are single-layered.

  • Beneath the palisade layer is the spongy layer. The cells of the spongy layer are more rounded and not so tightly packed. There are large air spaces between the cells. These cells contain fewer chloroplasts than those of the palisade layer. The pores or stomata of the epidermis open into chambers, which are connected to the air spaces between the spongy layer cells.

Plants with leaves all year round are evergreens, and those that shed their leaves are deciduous. Deciduous trees and shrubs generally lose their leaves in autumn. Before this happens, the leaves change colour. The leaves will grow back in spring.
Veins

Branching veins on underside of taro leaf


The 'veins' are a dense network of xylem, which supply water for photosynthesis, and phloem, which remove the sugars produced by photosynthesis. The pattern of the veins is called 'venation'.
In angiosperms the pattern of venation differs in the two main groups. Venation is usually is parallel in monocotyledons, but is an interconnecting network in broad-leaved plants (dicotyledons).
Hairs

Sticky trichomes of a carnivorous plantDrosera capensis with a trapped insect
Many leaves are covered in trichomes (small hairs) which have a wide range of structures and functions. Some trichomes are prickles, some are scaled, some secrete substances such as oil. Carnivorous plants secrete digestive enzymes from trichomes.
Waxy Cuticle The waxy cuticle is the waterproof, transparent outer surface of the leaf. It is waterproof to reduce water loss (transpiration) and transparent to allow light to enter the palisade cell.
Shape
What leaves look like on the plant varies greatly. Closely related plants have the same kind of leaves because they have all descended from a common ancestor. The terms for describing leaf shape and pattern is shown, in illustrated form, at Wikibooks.
Basic type

Leaves of the White Spruce (Picea glauca) are needle-shaped and their arrangement is spiral

  • Lycophytes have microphyll leaves.[2]

  • Ferns have fronds

  • Conifer leaves are typically needle-, awl-, or scale-shaped

  • Angiosperm (flowering plant) leaves: the standard form includes stipules, a petiole, and a lamina

  • Sheath leaves (type found in most grasses)

  • Other specialized leaves (such as those of Nepenthes)


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